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Carpinus betulas (Hornbeam)
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Pollen records are present from the Ludhamian to the Cromerian interglacials and are backed by macrofossil fruit remains. Records are frequent in the Hoxnian and there are fruit identifications in areas where the pollen percentage increases. The fruit is very durable and therefore is a frequent macrofossil (Godwin, 1975). Hornbeam has been present in each interglacial but only at low frequencies; it did not become noticeable as a forest component until the later stage of the Holocene cycle, when it saw an explosive expansion in central and western Europe.
During the last glaciation it is unlikely that it would have survived past the north of the Alps, and would have been partially devastated in periods of intensive settlement due to the fertility of occupied habitats. One of the reasons why hornbeam is present today as one of the main components of oak woodlands is due to its ability to regenerate in comparatively short time scales.

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